The Toffees looked like taking all three points from north London after Phil Jagielka and Kevin Mirallas scored following Emmanuel Adebayor’s 35th second opener. But Sigurdsson, charged with the task of playing in the hole where Bale has been so effective this season, salvaged a point for the hosts by scoring a crucial goal three minutes from time. Press Association Gylfi Sigurdsson’s late strike prevented Tottenham from slipping to another defeat, but Spurs’ reliance on Gareth Bale was laid bare as they had to settle for a 2-2 draw against Everton at White Hart Lane. Despite finding the net twice, Tottenham sorely missed the scoring talents of Bale and the creative wizardry of Aaron Lennon. All the pre-match talk had been about how Spurs would struggle without the Welshman’s goals, but they gave the perfect response by breaking the deadlock with the second quickest goal of the season. In the first move of the match, Everton’s defence backed off Jan Vertonghen, he delivered a teasing ball across the box, and Adebayor stuck his right leg and prodded the ball past Tim Howard. Everton were shell-shocked by the goal, but it did not take them long to come back in to the game. Michael Dawson almost gifted them a 10th-minute equaliser when he passed the ball straight to Leon Osman 15 yards out, but he curled high and wide. Dawson then cut out Victor Anichebe’s cross as a row of Everton attackers lined up to convert, but unfortunately for the Tottenham captain, his defensive colleague Vertonghen let him down from the resulting corner. The Belgian lost Jagielka at the back post and he nodded Leighton Baines’ cross in to the bottom corner. Spurs thought they should have had a penalty on the half hour when Clint Dempsey’s cross brushed Darron Gibson’s arm, but Andre Marriner waved play on. Everton sounded an early warning just after the restart when Gibson struck a curling shot just wide, and with 52 minutes on the clock, they were ahead. Mirallas drifted in from the right flank and drifted past Scott Parker and Steven Caulker before slotting the ball beyond Hugo Lloris. Spurs went straight on the attack and almost scored an equaliser through Mousa Dembele, whose 20-yard shot ricocheted off John Heitinga’s foot and Howard’s arm on to the bar. Heitinga almost sealed the win at the other end soon after but Dawson put in a crunching goal-saving tackle to deny the Dutchman. Spurs’ hopes of an equaliser looked to have faded but with three minutes left, Adebayor’s shot came back off the post and Sigurdsson converted.
Linking extreme interannual changes in prey availability to foraging behaviour and breeding investment in a marine predator, the macaroni penguin
Understanding the mechanisms that link prey availability to predator behaviour and population change is central to projecting how a species may respond to future environmental pressures. We documented the behavioural responses and breeding investment of macaroni penguins Eudyptes chrysolophus across five breeding seasons where local prey density changed by five-fold; from very low to highly abundant. When prey availability was low, foraging trips were significantly longer and extended overnight. Birds also foraged farther from the colony, potentially in order to reach more distant foraging grounds and allow for increased search times. These extended foraging trips were also linked to a marked decrease in fledgling weights, most likely associated with reduced rates of provisioning. Furthermore, by comparing our results with previous work on this population, it appears that lowered first-year survival rates associated, at least partially, with fledging masses were also evident for this cohort. This study integrates a unique set of prey density, predator behaviour and predator breeding investment data to highlight a possible behavioural mechanism linking perturbations in prey availability to population demography.